Catocala from Pennsylvania

The following checklist for Ohio was sent to me by Jeff Crolla. The list was compiled in 1992 (Rings et al.).

I believe most of the Ohio sepcies are also found in Pennsylvania. Over time, I will refine the list and add thumbnails. There may be some erroneous inclusions for Pennsylvania, and doubtless there are some omissions.

* 8787 agrippina Strecker, 1874 Agrippina Underwing

**8878 Catocala alabamae; Alabama Underwing; 30-40mm; yes; Uniform greenish-grey forewing may have thin but dark (contrasting) medial lines or they may be nearly absent (titania); alabamae often displays dash in median/anal angle area, missing or nearly absent in titania. Brown shading between post medial line and subterminal line. Hindwing inner black band thin, outer band thick, truncated before being followed by large black dot. Early flight season.

* 8834 amatrix (Hübner, [1813]) Sweetheart Underwing

* 8878 amica (Hubner, 1818) Girlfriend Underwing

* 8849 andromedae (Guenee, 1852) Andromeda Underwing

* 8775 antinympha (Hübner, [1823]) Sweetfern Underwing

* 8777 badia Grote and Robinson, 1866 Bay Underwing

** 8867 Catocala blandula; Charming Underwing, wingspan: 40-50mm.
Pale basal area of fw distinguishes mira from blandula (dark brown) and crataegi (black). Fw has dark contrasting lines as in crataegi. There is also considerable brown in subterminal area and subreniform spot is very conspicuous and usually brown. A light area runs obliquely from the costa to the subreniform spot. hw is deep orange and has a complete inner black band. The outer black band is unbroken. Lynn Scott image.

* 8817 briseis W.H. Edwards, 1864, Breseis Underwing

** 8832 Catocala cara; Darling; 70-85mm. Note deep maroon almost purple cast to fws, mixed with very pale green. Lower half of am & pm lines is barely visible. There are no distinctive bars or dashes. The two upper "teeth" on the pm line are thin and long.
The hindwing bands are pink. There is heavy black checking on the off-white hindwing fringe. The relatively thick black median band of the hindwing almost reaches the inner margin which is usually heavily adorned with dark hairs.
Tim Dyson image.

** 8802 Catocala cerogama; Yellow-Banded; 70-80mm. Several different forms. Pm line distinct, dark and has elongate pair of projections. Pm and am lines meet inner margin in relative proximity. Closed subreniform spot lighter than surrounding areas, shaped a bit like an arrowhead with point toward body. Hws are distinctive. Jean-Benoît Duval image.

** 8802 Catocala cerogama; Yellow-Banded; 70-80mm.

Form ruperti: Forewings are almost uniform grey-brown with white scales outlining subterminal line.

The hindwings are distinctive. Tim Dyson image.

** 8802 Catocala cerogama; Yellow-Banded; 70-80mm.

Form bunkeri: Forewings are dark in median and basal areas. Hw golden band is reduced in thickness. Basal hairs are brown instead of yellow.

The hindwings are distinctive. Tim Dyson image.

* 8872 clintoni Grote, 1864 Clinton's Underwing

** 8851 Catocala coccinata; Scarlet; wingspan: 57-70mm: There are usually diffuse basal and anal dashes on an otherwise light grey, mottled fw. Hw fringe is white (often with some salmon scaling) and is heavily checked. I find "tooth" just below pair of very elongated "teeth" is much reduced and is quite rounded, usually allowing considerable room for lighter patch of scales. Dark bar crosses thorax. Reniform spot tends be to light, often with greenish cast. Joe Garris image.

** 8833 concumbens Sleepy Underwing or Pink Underwing, 60-75mm). This brown thoracic collar is quite evident in this image as is the interruption in the pm line by the open subreniform spot.

The white hindwing fringe is only lightly checked on the wing veins. The vibrant pink bans are distinct in colour and also in their relatively smooth contour.

* 8772 consors (J.E. Smith, 1797) Consort Underwing

** 8858 Catocala crataegi; Hawthorn Underwing, wingspan: 40-50mm Determination is based on dark (black) shading in forewing basal area continuing along inner margin to anal angle and brown shading beyond postmedial line. Definite greenish cast to median area. Lower wing has outer black band, broken near anal angle, distinguishing crataegi from blandula.

dulciola; Quiet/Sweet Underwing, wingspan: 40-45mm;

A double, black, almost smooth antemedial line (inner line broken only near the costa) with other lines obscure distinguishes dulciola. There is also a dark basal dash on an otherwise plain, light grey forewing.

** 8772 gracilis; Graceful Underwing, wingspan 40-45mm.

Gracilis closely resembles sordida but the hindwing loop is not complete in gracilis as it is in sordida. Gracilis also tends to have a darker inner margin and there is frequently (99% of the time, Dale Schweitzer, via Joe Garris), but not always, a basal dash. Sordida never has the basal dash.

* 8778 habilis Grote, 1872 HABILIS UNDERWING

* 8840 illecta Walker, [1858] Magdalen Underwing

* 8791 insolabilis Guenee, 1852 Inconsolable Underwing

* 8829 junctura Walker, [1858]Joined Underwing

* 8794 lacrymosa Guenee, 1852 Tearful Underwing

* 8878.1 lineella Grote, 1872 Little Lined Underwing

* 8788.1 luctuosa Hulst, 1884 Yellow-grey Underwing

* 8793 maestosa Hulst, 1884 Sad Underwing

* 8804 marmorata Edwards, 1864 Marbled Underwing


Little Yellow-Orange Underwings: Wingspans: 35-56mm
These tend to be slightly larger on average than preceding group.

** 8876 Catocala micronympha, the Little Nymph Underwing, (wingspan: 35-50mm). John Himmelman image.
The usual specimens have grey forewings shaded with green, brown, black and white tints. There is usually a darkened band passing from the costa through the reniform spot to the outer margin.

There is high variability with this species. Full size Joe Garris photo of C. micronympha form hero.

** 8877 connubialis, Connubial Underwing, (wingspan: 35-50mm). The forewing is highly variable and there are several different forms: "sancta" Hulst is the typical form and has forewings with white ground colour and sharply contrasting black lines and markings. There is also brown shading between post medial and subterminal lines; "cordelia" H. Edwards is coloured as above but markings are faint; "pulverulenta" Brower has grey-green forewings with faint markings, sometimes none; "broweri" Muller is the melanic form with forewings that are dark green, almost black. The hindwing has a separate anal spot and the outer band ends with a straight cut. Leroy Simon image.

** 8864 Catocala grynea ; Woody Underwing, wingspan: 40-50mm

The forewing is a dull greenish grey with orangey-brown shading along the inner margin. The antemedial, median and postmedial lines are quite faint.

Praeclara is somewhat similar but has a break in the brown shading between the am and pm lines. Praeclara also has a paler orange hindwing color.
Tim Dyson image.

** 8874 Catocala minuta; Little Underwing; 35-45mm; Fw generally grey-brown with greatly widened (near costa only), white, stl. Hw outer black band unbroken, inner band in complete loop, considerable brown scaling along im. Forms: "eureka" Schwarz: blackish area between lines; "hiseri" Cassino: dull grey fw, very faint markings; "mellitula" Hulst: blackish basal patch from costa to im; "obliterata" Schwarz: melanic form, almost completely black fw; "parvula" W. H. Edwards: broad dark patch along fw im.


Solid Black Underwings: Smallest to Largest, Similar Species Paired

** 8781 judith, Judith's Underwing, (wingspan: 45-55mm), is one of the smaller "black" underwings.

The forewing is a uniform light grey with thin and only slightly darkened antemedial, median and postmedial lines. There are no darkened dashes (slight anal dash) or transverse lines. The reniform area is slightly darkened while the area just before the subterminal line is a bit lighter. Note absence of hw white fringe. Joe Garris photo.

** 8782 Catocala flebilis, yes; Mournful, wingspan: 54-65mm. Diffuse black band runs from basal area to outer margin just below apex, interrupted by pale grey subreniform spot. Reniform spot is filled with brown; there is additional brown outside postmedial line. No anal dash as there is in angusi. The hindwings have white fringe. J. K. Adams image.

** 8773 Catocala epione , yes; Epione Underwing, wingspan: 55-65mm

The pm line is squared and has a brown band and then a light band just outside the line.

The hindwing is black with pure white fringe with no barring.
John Himmelman image.

Catocala residua, Residua Underwing, yes, (wingspan: 58-73mm), Distinguished from C. obscura by grey hindwing fringe of residua. C. obscura has white to off-white fringe, tends to be less common in northern portions of its range. Fw subterminal line/area of C. residua tends to be pale as is its open, elongated subreniform spot.
Devoid of significant markings except for hint of dark bar running from basal area through reniform spot to om just below apex.

Catocala dejecta; Dejected; wingspan 56-73mm. Pale grey patch from reniform and subreniform juncture to costal margin is diagnostic. Note open, light "pork chop" shape of subreniform spot. Two "teeth" in pml above subreniform spot are relatively short and blunt. Upper portion of aml thick and black; lower half has light rounded lobes outlined in black. Hw fringe is white with black wing veins extending like "teeth".

** 8783 Catocala angusi WO; Angus' Underwing, 60-74mm.

Dark dashes/streaks in basal & anal areas. Reniform spot: light brown filling. Hw fringe black except for white region at apex. James K. Adams image.

** 8783 Catocala angusi WO; Angus' Underwing, 60-74mm; Dark dashes/streaks in basal & anal areas. Reniform spot: light brown filling. Hw fringe black except for white region at apex. Form lucetta has broad black band from basal area to om, broken only by reniform & subreniform spots. James K. Adams image.

** 8792 Catocala vidua, yes; Widow; wingspan: 70-80mm
The ground colour of the forewing is light grey. There is a distinguished dark arc running through the top of the reniform spot to just below the apex. There are heavy, dark anal and basal dashes, connecting to a dark median bar, running parallel to the inner margin.
The am line is thick and black in its upper half.
The reniform spot consists of two almost concentric irregular ovals. The subreniform spot is light, open, but constricted as it meets the pm line. The hindwing is black with broad, white fringe, only lightly interrupted. Marie Winn image.

** 8803 Catocala relicta ; Forsaken, White, Relict; 70-80mm: Considerable variation with regard to black/white concentrations on fws.
Typical specimens have basal and subterminal areas with blackish scales.

Black hws, with brilliant even white inner band and white fringe, are distinctive. June until October.

** 8803 Catocala relicta ; Forsaken, White, Relict; 70-80mm: Considerable variation with regard to black/white concentrations on fws. Form clara: basal and subterminal areas predominantly white.

Typical specimens have basal and subterminal areas with blackish scales. Black hws, with brilliant even white inner band and white fringe, are distinctive. June until October.

** 8803 Catocala relicta ; Forsaken, White, Relict; 70-80mm: Considerable variation with regard to black/white concentrations on fws. Form phrynia: evenly dusted with grey over entire forewing. Typical specimens have basal and subterminal areas with blackish scales. Black hws, with brilliant even white inner band and white fringe, are distinctive. June until October, poplars and willows


Large Orange-Salmon Underwings: Wingspans: 65mm, usually 70-95mm

** 8822 meskei; Meske's Underwing; wingspan: 65-75mm
The forewings are less distinctly marked compared to unijuga and there is some red-orange at the hindwing apex and just inside the fringe along the outer margin in meskei that is lacking in unijuga. The forewing subreniform spot is opened or connected to the postmedian line. The hindwing postmedian band is nearly straight, turned in and tapering near the anal angle. Tim Dyson image.

* 8863 mira Grote, 1876 Wonderful Underwing

* 8855 miranda Edwards, 1881 Miranda Underwing

* 8796 nebulosa Edwards, 1864 Clouded Underwing

* 8798 neogama (J.E.Smith, 1797) The Bride Underwing

* 8784 obscura TL Strecker, 1873 Obscure Underwing

* 8795 palaeogama Guenee, 1852 Oldwife Underwing

* 8806 parta Guenee, 1852 Mother Underwing

** 8771 Catocala piatrix ; Penitent, wingspan: 68-84mm. Fw: light-colored band/bar extending from light coloured, triangular subreniform spot along the am line to the costa. Brown, kidney-shaped reniform spot surrounded in pale grey, outlined in black, usually with distinct black along wing veins in a dark area outside the reniform spot.

Hw: fringe lightly barred;s lighter in color than deeper orange on rest of wing.

** 8865 praeclara; Praeclara Underwing, (40-50mm); The forewing is a pale greenish-grey with considerable contrasting brown shading beyond post medial line.

There is a black basal dash very close to the inner margin and another dash, higher up in the median area.

Note the contrasting shapes of the reniform and subreniform spots. Tim Dyson image.

Catocala pretiosa Precious Underwing; 40-50mm; This species is quite similar to crataegi and mira, but pretiosa has considerable creamy white colouration in the median area. The lower portion of the basal area is also light, not dark as in crataegi. The inner black band of the hindwing forms a loop and the outer band is broken (not complete as in mira, followed by a dot that tapers to the anal angle.

*8788 retecta Grote, 1872 Underwing Moth

* 8780 robinsoni Grote, 1872 Robinson's Underwing

** 8779 serena Serene Underwing. The head and collar and abdomen are brown while the thorax is grey. The am and pm lines are thin but very dark and distinct on an otherwise drab, almost uniformly olive-grey forewing. The black marginal band of the lower wings is indented at the center, and the relatively narrow, deep yellow median band parallels this indentation. Basal hairs are brown. Catocala serena courtesy of Carroll Rudy, July 28, Calumet County, Wisconsin.

* 8873 similis Edwards, 1864 Similar Underwing

* 8846 sordida Grote, 1877 Sordid Underwing

* 8797 subnata Grote, 1864 Youthful Underwing

* 8789 ulalume Strecker, 1878 Ulalume Underwing

** 8801 Catocala ilia; Ilia; wingspan: 65-82mm. Several different forms, most have characteristic white area in and around reniform spot. Diffuse dark arc running from this spot to just below apex. Subreniform spot squarish, concave inner and outer edges and elongated constriction connecting it to pml. White dots near fw om in character with the overall "contrasting" appearance.

** 8801 Catocala ilia; Ilia; form conspicua

In this form the entire reniform spot is heavily suffused with white scaling on an otherwise darker ground colour. Hence the form name "conspicua".

Tim Dyson image.

** 8801 Catocala ilia; Ilia; form satanas

In this melanic form the entire forewing, including the reniform spot is very dark. Hence the form name "satanas".

The dark basal streak is still evident on this form.

Tim Dyson image.

** 8801 Catocala ilia; Ilia; form normani

In this semi-melanic form the entire forewing, excluding the reniform spot, is relatively dark. The brownish, kidney-shaped center of the reniform spot is outlined in white.

The basal streak and subapical arc are still visible.

Tim Dyson image.

** 8770 Catocala innubens; Betrothed; 55-72mm. Forewing is mottled with white, grey and brown, and subrenifrom spot tends to be lighter in colour, although it is sometimes obscured by an indistinct blackish bar which runs from middle of basal/thorax connection to just below much lighter apex at outer margin.

Jim Vargo image.

** 8770 Catocala innubens form scintillans; Betrothed; 55-72mm.

In form scintillans most of the basal area (all but lower third) and all of the median area of the forewing is dark reddish brown. The area outside the pm line is grey.

Vernon A Brou image.

** 8857.1 Catocala umbrosa; mm; Double brown am line, inward line fainter, filled with off white; reniform spot brown center outlined in black , off white and black again; subreniform, large, pale brown, closed. Pm line wit htwo elongated upper teeth, next tooth reduced, next two progressively longer, rounded, final lobe rounded and shorter. HW with dark scaling/hairs in basal median area along im

Catocala muliercula; Little Wife Underwing; Dark, distinct upper half am line runs obliquely toward anal angle, turns in two lobes to im; basal area dark brown; inner brown kidney shaped reniform spot, oulined in black then light then black again; open, lighter subreniform spot with light area from it to costa; median area greyish-blue-brown; subterminal area brown; terminal area greyish-blue-brown; lighter patch at apex. Hw dark bands thick, much dark scaling in basal area covering 2/3, joining inner black band to im.

** 8857 Catocala ultronia; Ultronia Underwing, wingspan: 50-63mm.

Fws typically gray-brown, with a distinct and very dark inner margin and characteristic light brown patch, underscored by very dark arc, near wingtip. Underwings can be yellow to orange to salmon.

** 8857 Catocala ultronia form lucinda; Ultronia Underwing, wingspan: 50-63mm.

In form lucinda most of the forewing is bright grey.

On all forms there is extensive orange-salmon colouration on hw ventral surface, and there is a dark discal lunule.

Catocala ultronia, form nigrescens, the Ultronia Underwing, wingspan: 50-63mm. In the melanic form nigrescens, the dorsal forewing is very dark. Even darker subapical arc, basal dash and dash near anal angle are still visible. Dark basal hairs on hindwing. Harold J. Vermes slide, used with permission from his son.

Catocala ultronia form celia.

In form celia there is a wide, light grey band separating a dark region along inner margin and a dark patch near the apex. The ventral surface of forewings of all forms has a generous suffusion of orange-salmon scales in the lower half of the median area.

* 8805 unijuga Walker, [1858] Once-married Underwing

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